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COL6A3 – Ullrich congenital muscular dystrophy

COL6A3 encodes the α3 chain of type VI collagen, a critical component of the extracellular matrix in muscle and skin. Ullrich congenital muscular dystrophy (UCMD; MONDO:0000355) is characterized by early-onset muscle weakness, proximal joint contractures, distal joint hyperextensibility, and progressive respiratory compromise. Initial linkage and sequencing studies in a consanguineous family with three affected siblings established autosomal recessive inheritance of UCMD due to homozygous COL6A3 splice-donor mutations, demonstrating exon skipping, reduced collagen VI deposition, and an intermediate phenotype (PMID:11992252).

Subsequent case reports expanded the mutation spectrum to include one patient with a homozygous nonsense mutation (c.1393C>T (p.Arg465Ter)) and another with homozygous nonsense (c.7024C>T (p.Arg2342Ter)), both causing absent collagen VI in muscle and severe UCMD (PMID:11992252). More recently, de novo heterozygous splice-site variants (c.6210+1G>A) were identified in classical UCMD presentations, indicating autosomal dominant mechanisms can also underlie UCMD (PMID:35832501).

In multi-patient studies, immunohistochemical analysis of 15 UCMD patients revealed collagen VI deficiency linked to COL6A3 in 40% of cases by haplotype and linkage, underscoring genetic heterogeneity yet common involvement of COL6A3 in UCMD (PMID:12011280). Targeted NGS in cohorts confirmed COL6A3 pathogenic variants in both recessive and dominant contexts, accounting for a significant fraction of UCMD cases and highlighting variable expressivity.

Inheritance of COL6A3-related UCMD is predominantly autosomal recessive, but de novo dominant splice mutations have been documented. To date, at least seven unrelated probands harbor three loss-of-function (two nonsense, one in-frame pseudoexon) and two splice-site variants, with segregation in multiple families and de novo occurrence. A representative pathogenic variant is c.7024C>T (p.Arg2342Ter), which truncates the collagen VI α3 chain and abolishes matrix assembly.

Functional assays have confirmed the pathogenicity of COL6A3 variants: splice-site and nonsense mutations lead to intracellular misfolding, degradation of mutant α3(VI) domains, loss of extracellular collagen VI network integrity, and impaired filament organization in patient fibroblasts (PMID:15965965; PMID:11992252). These defects mirror the clinical phenotype of UCMD, supporting a loss-of-function mechanism.

Collectively, the genetic and experimental data support a Strong clinical validity for the COL6A3–UCMD association. COL6A3 mutations explain a substantial subset of UCMD cases, enabling accurate diagnosis, genetic counseling, and informing emerging splice-modulating therapies. Key take-home: COL6A3 pathogenic variants cause UCMD via recessive and de novo dominant mechanisms, leading to definitive collagen VI deficiency and a clinically actionable genotype–phenotype correlation.

References

  • American journal of human genetics • 2002 • Mutations in COL6A3 cause severe and mild phenotypes of Ullrich congenital muscular dystrophy. PMID:11992252
  • Journal of cellular physiology • 2006 • Ultrastructural defects of collagen VI filaments in an Ullrich syndrome patient with loss of the alpha3(VI) N10-N7 domains. PMID:15965965
  • Acta myologica • 2022 • Autosomal dominant Ullrich congenital muscular dystrophy due to a de novo mutation in COL6A3 gene. A case report. PMID:35832501

Evidence Based Scoring (AI generated)

Gene–Disease Association

Strong

Seven unrelated probands (five recessive, two de novo dominant), segregation in consanguineous family, concordant functional data

Genetic Evidence

Moderate

Seven pathogenic COL6A3 variants (three loss-of-function, two splice-site) identified in multiple families including de novo occurrences

Functional Evidence

Moderate

Mutant α3(VI) misfolding and assembly failure in cell and tissue assays, consistent with human UCMD phenotype