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CNNM2 – Hypomagnesemia, Seizures, and Intellectual Disability 1

Hypomagnesemia, seizures, and intellectual disability 1 (HOMGSMR1) is a rare autosomal recessive neurodevelopmental disorder caused by biallelic or de novo heterozygous mutations in CNNM2 that disrupt magnesium homeostasis (PMID:24699222). Initial reports described five families with recessive CNNM2 variants presenting with severe hypomagnesemia, brain malformations, intellectual disability and seizures, establishing a key genetic link for this syndrome (PMID:24699222).

Subsequent cohort studies expanded the spectrum to 11 sporadic patients carrying heterozygous CNNM2 mutations, confirming loss-of-function effects in Mg2+ uptake assays and demonstrating decreased plasma membrane expression of mutant proteins (PMID:33600043). Additional de novo heterozygous missense variants were reported in individual cases presenting with refractory epilepsy or mild intellectual disability, sometimes in absence of overt hypomagnesemia (PMID:34490037; PMID:35846113; PMID:35806288). A 2024 analysis of seven novel heterozygous variants further underscored phenotypic variability, with some patients lacking neurological features despite pronounced renal Mg2+ wasting (PMID:38519529). Most recently, two unrelated families with novel p.Glu298del and p.Pro360Arg variants recapitulated the core HSMR phenotype and confirmed impaired CNNM2 membrane localization and Mg2+ efflux (PMID:40612795).

The CNNM2 variant spectrum encompasses missense changes (e.g., c.1079C>G (p.Pro360Arg)), frameshifts (e.g., c.274del (p.Ala92fs)) and nonsense alleles (e.g., c.12T>A (p.Cys4Ter)), as well as small in-frame deletions (e.g., p.Glu298del) detected in both homozygous and heterozygous states across diverse populations. No clear founder alleles have emerged, but recurrent CBS-domain mutations (e.g., p.Thr568Ile) highlight critical functional hotspots (PMID:25184538).

Functional characterization has integrated membrane topology mapping in kidney cells, revealing an extracellular N-terminus and intracellular C-terminus with glycosylation at Asn-112 enhancing surface stability (PMID:22399287), in vitro Mg2+ uptake assays confirming loss of transporter activity and TRPM7 regulation, and zebrafish cnnm2 knockdown models demonstrating neurodevelopmental rescue by wild-type but not mutant cRNA (PMID:24699222). Crystal structures of the CBS module have further shown that nucleotide binding and pathogenic mutations (e.g., T568I) induce conformational locking that may underlie defective Mg2+ sensing (PMID:25184538).

Collectively, clinical, genetic and experimental data support a Strong gene–disease association: over 21 independent probands, segregation in seven families, and concordant functional evidence demonstrating a loss-of-function mechanism. No credible conflicting evidence has emerged to date. Additional reports continue to refine genotype-phenotype correlations and may exceed current ClinGen scoring maxima.

Key Take-home: CNNM2 mutation testing enables diagnosis of HOMGSMR1 in patients with hypomagnesemia, seizures, and intellectual disability and guides genetic counseling and potential Mg2+ supplementation strategies.

References

  • The Journal of biological chemistry • 2012 • Membrane topology and intracellular processing of cyclin M2 (CNNM2). PMID:22399287
  • PLoS genetics • 2014 • CNNM2 mutations cause impaired brain development and seizures in patients with hypomagnesemia. PMID:24699222
  • The Biochemical journal • 2014 • Nucleotide binding triggers a conformational change of the CBS module of the magnesium transporter CNNM2 from a twisted towards a flat structure. PMID:25184538
  • Human mutation • 2021 • The phenotypic and genetic spectrum of patients with heterozygous mutations in cyclin M2 (CNNM2). PMID:33600043
  • Frontiers in genetics • 2021 • Case Report: CNNM2 Mutations Cause Damaged Brain Development and Intractable Epilepsy in a Patient Without Hypomagnesemia. PMID:34490037
  • International journal of molecular sciences • 2022 • The p.Pro482Ala Variant in the CNNM2 Gene Causes Severe Hypomagnesemia Amenable to Treatment with Spironolactone. PMID:35806288
  • Frontiers in genetics • 2022 • Novel CNNM2 Mutation Responsible for Autosomal-Dominant Hypomagnesemia With Seizure. PMID:35846113
  • Scientific reports • 2024 • Hypomagnesaemia with varying degrees of extrarenal symptoms as a consequence of heterozygous CNNM2 variants. PMID:38519529
  • Frontiers in genetics • 2025 • Two novel variants in CNNM2 disrupts magnesium efflux leading to neurodevelopmental disorders. PMID:40612795

Evidence Based Scoring (AI generated)

Gene–Disease Association

Strong

21 probands across 7 families, multi-family segregation and concordant functional data

Genetic Evidence

Strong

21 probands with recessive and de novo variants, heterogeneous variant classes reaching genetic cap

Functional Evidence

Moderate

Multiple in vitro topology, stability and Mg2+ transport assays, zebrafish knockdown and rescue models demonstrate loss-of-function